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For example, the GRX [2Fe–2S] clusters form complexes with BOLA proteins, in which the [2Fe–2S] cluster is ligated using the GRX conserved Cys, a Cys from GSH, and His or Cys residues on the BOLA protein

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373 In fact, when BMSCs are exposed to exendin-4, the expression of genes related to bone development factors such as Runx and Osterix, as well as genes responsible for producing the bone matrix such as Balp and Bglap, is upregulated